:root {
  color-scheme: light;
  --paper: #f7f9f3;
  --surface: #fbfcf8;
  --surface-strong: #eef5eb;
  --ink: #10231f;
  --muted: #5f6f69;
  --line: #dce8de;
  --accent: #119e92;
  --accent-dark: #0b6f67;
  --accent-soft: #dff3ed;
  --cool-soft: #edf4f8;
  --shadow: 0 24px 70px rgba(34, 73, 65, 0.12);
  --radius: 8px;
  --header-height: 72px;
}

* {
  box-sizing: border-box;
}

html {
  scroll-behavior: smooth;
}

body {
  margin: 0;
  background: var(--paper);
  color: var(--ink);
  font-family: ui-sans-serif, system-ui, -apple-system, BlinkMacSystemFont, "Segoe UI", sans-serif;
  line-height: 1.5;
  text-rendering: optimizeLegibility;
}

body.menu-open {
  overflow: hidden;
}

img {
  display: block;
  max-width: 100%;
  height: auto;
}

a {
  color: inherit;
  text-decoration: none;
}

p {
  margin: 0;
}

.section-shell {
  width: min(1180px, calc(100% - 40px));
  margin: 0 auto;
}

section[id] {
  scroll-margin-top: 96px;
}

.site-header {
  position: sticky;
  top: 0;
  z-index: 20;
  height: var(--header-height);
  display: grid;
  grid-template-columns: auto 1fr;
  align-items: center;
  gap: 24px;
  padding: 0 28px;
  border-bottom: 1px solid rgba(220, 232, 222, 0.82);
  background: rgba(247, 249, 243, 0.9);
  backdrop-filter: blur(18px);
  -webkit-backdrop-filter: blur(18px);
}

.brand {
  display: inline-flex;
  align-items: center;
  gap: 12px;
  min-width: 0;
}

.brand img {
  width: 44px;
  height: 44px;
  border-radius: var(--radius);
  object-fit: cover;
  box-shadow: 0 8px 18px rgba(16, 35, 31, 0.12);
}

.brand strong,
.brand small {
  display: block;
  line-height: 1.1;
  white-space: nowrap;
}

.brand strong {
  font-size: 0.96rem;
  letter-spacing: 0.02em;
}

.brand small {
  margin-top: 4px;
  color: var(--muted);
  font-size: 0.78rem;
}

.site-nav {
  justify-self: end;
  display: flex;
  align-items: center;
  gap: 6px;
}

.site-nav a {
  min-height: 40px;
  display: inline-flex;
  align-items: center;
  padding: 0 14px;
  border-radius: 999px;
  color: #2a403a;
  font-size: 0.94rem;
  transition: background 180ms ease, color 180ms ease, transform 180ms ease;
}

.site-nav a:hover,
.site-nav a:focus-visible,
.site-nav a.is-active,
.site-nav a[aria-current="page"] {
  background: var(--accent-soft);
  color: var(--accent-dark);
}

.site-nav a:active,
.button:active {
  transform: translateY(1px);
}

.menu-button {
  display: none;
  justify-self: end;
  border: 1px solid var(--line);
  border-radius: 999px;
  min-height: 42px;
  padding: 0 18px;
  background: var(--surface);
  color: var(--ink);
  font: inherit;
}

.hero {
  position: relative;
  overflow: hidden;
  min-height: min(690px, calc(100dvh - var(--header-height) - 56px));
  display: flex;
  align-items: center;
  border-bottom: 1px solid var(--line);
  background:
    linear-gradient(90deg, rgba(247, 249, 243, 0.98) 0%, rgba(247, 249, 243, 0.86) 42%, rgba(247, 249, 243, 0.58) 100%),
    url("assets/images/hero-evo-devo-background.png") center right / cover no-repeat;
}

.hero::after {
  content: "";
  position: absolute;
  inset: 0;
  pointer-events: none;
  background:
    radial-gradient(circle at 18% 28%, rgba(255, 255, 255, 0.72), transparent 34%),
    linear-gradient(180deg, transparent 70%, rgba(247, 249, 243, 0.8) 100%);
}

.hero-grid {
  position: relative;
  z-index: 1;
  display: grid;
  grid-template-columns: minmax(0, 0.88fr) minmax(520px, 0.92fr);
  align-items: center;
  gap: 56px;
  padding-top: 76px;
  padding-bottom: 84px;
}

.hero h1 {
  max-width: 760px;
  margin: 0;
  font-size: 4.2rem;
  line-height: 0.98;
  letter-spacing: 0;
}

.button {
  min-height: 46px;
  display: inline-flex;
  align-items: center;
  justify-content: center;
  border-radius: 999px;
  padding: 0 22px;
  font-weight: 700;
  white-space: nowrap;
  transition: transform 180ms ease, background 180ms ease, border-color 180ms ease;
}

.button.primary {
  background: var(--accent-dark);
  color: #f7fbf7;
}

.button.primary:hover,
.button.primary:focus-visible {
  background: #084f4b;
}

.button.secondary {
  border: 1px solid var(--line);
  background: var(--surface);
  color: var(--ink);
}

.button.secondary:hover,
.button.secondary:focus-visible {
  border-color: var(--accent);
  background: var(--accent-soft);
}

.hero-brief {
  max-width: 650px;
  display: grid;
  gap: 14px;
  padding: 30px;
  border: 1px solid rgba(220, 232, 222, 0.56);
  border-radius: var(--radius);
  background: rgba(251, 252, 248, 0.36);
  backdrop-filter: blur(8px) saturate(120%);
  -webkit-backdrop-filter: blur(8px) saturate(120%);
  box-shadow: 0 24px 72px rgba(34, 73, 65, 0.08);
}

.hero-brief p {
  color: #41564f;
  font-size: 1.04rem;
  line-height: 1.58;
}

.intro-band {
  border-top: 1px solid var(--line);
  border-bottom: 1px solid var(--line);
  background: var(--surface);
}

.intro-grid {
  display: grid;
  grid-template-columns: 0.9fr 1.1fr;
  gap: 56px;
  padding: 70px 0;
}

.intro-lede {
  font-size: 2rem;
  line-height: 1.12;
  font-weight: 750;
  letter-spacing: 0;
}

.intro-copy {
  display: grid;
  gap: 18px;
  color: var(--muted);
  font-size: 1.02rem;
}

.research-section,
.people-section,
.contact-section {
  padding: 104px 0;
}

.section-heading {
  display: grid;
  gap: 16px;
  max-width: 720px;
  margin-bottom: 36px;
}

.section-heading h2,
.platform-grid h2,
.contact-card h2 {
  margin: 0;
  font-size: 3rem;
  line-height: 1.05;
  letter-spacing: 0;
}

.section-heading p,
.platform-grid p,
.contact-card > div > p {
  color: var(--muted);
  font-size: 1.06rem;
}

.page-main {
  min-height: calc(100dvh - var(--header-height));
}

.page-hero {
  position: relative;
  overflow: hidden;
  padding: 104px 0 78px;
  border-bottom: 1px solid var(--line);
  background:
    linear-gradient(90deg, rgba(247, 249, 243, 0.98), rgba(247, 249, 243, 0.78)),
    url("assets/images/hero-evo-devo-background.png") center right / cover no-repeat;
}

.page-hero::after {
  content: "";
  position: absolute;
  inset: 0;
  pointer-events: none;
  background:
    radial-gradient(circle at 16% 18%, rgba(255, 255, 255, 0.66), transparent 30%),
    linear-gradient(180deg, transparent 58%, rgba(247, 249, 243, 0.9));
}

.page-hero .section-shell {
  position: relative;
  z-index: 1;
  display: grid;
  gap: 18px;
}

.page-hero h1 {
  max-width: 760px;
  margin: 0;
  font-size: 4.4rem;
  line-height: 1;
  letter-spacing: 0;
}

.page-hero p {
  max-width: 780px;
  color: var(--muted);
  font-size: 1.12rem;
}

.research-grid {
  display: grid;
  grid-template-columns: minmax(0, 1.35fr) minmax(340px, 0.85fr);
  gap: 18px;
}

.research-list {
  display: grid;
  gap: 18px;
}

.research-row {
  display: grid;
  grid-template-columns: 390px minmax(0, 1fr);
  gap: 34px;
  align-items: center;
  overflow: hidden;
  padding: 22px;
  border: 1px solid var(--line);
  border-radius: var(--radius);
  background: rgba(251, 252, 248, 0.76);
  box-shadow: 0 18px 54px rgba(34, 73, 65, 0.07);
}

.research-row:nth-child(2) {
  background: rgba(223, 243, 237, 0.52);
}

.research-row:nth-child(3) {
  background: rgba(237, 244, 248, 0.72);
}

.research-row img {
  width: 100%;
  aspect-ratio: 16 / 10;
  object-fit: contain;
  padding: 8px;
  border-radius: calc(var(--radius) - 2px);
  background: rgba(251, 252, 248, 0.68);
}

.research-row div {
  display: grid;
  gap: 12px;
}

.research-card {
  display: grid;
  gap: 18px;
  align-content: start;
  overflow: hidden;
  border: 1px solid var(--line);
  border-radius: var(--radius);
  background: var(--surface);
}

.research-card-large {
  grid-row: span 2;
}

.research-card.tinted {
  background: var(--accent-soft);
}

.research-card.cool {
  background: var(--cool-soft);
}

.research-card img {
  width: 100%;
  aspect-ratio: 16 / 9;
  object-fit: cover;
  background: var(--surface);
}

.research-card-large img {
  aspect-ratio: 16 / 10;
}

.research-card div {
  display: grid;
  gap: 12px;
  padding: 0 22px 24px;
}

.research-card h3,
.research-row h3,
.pi-feature h2,
.pi-feature h3,
.person-card h3,
.publication-card h3,
.publication-row h2,
.contact-details h3 {
  margin: 0;
  font-size: 1.28rem;
  line-height: 1.18;
  letter-spacing: 0;
}

.research-card p,
.research-row p,
.person-card p,
.person-details-body p,
.publication-card p,
.publication-body p,
.contact-details p,
address {
  color: var(--muted);
}

.platform-section {
  padding: 88px 0;
  background: var(--surface-strong);
}

.platform-grid {
  display: grid;
  grid-template-columns: minmax(0, 0.9fr) minmax(420px, 1fr);
  gap: 56px;
  align-items: start;
}

.platform-grid > div:first-child {
  display: grid;
  gap: 18px;
}

.platform-list {
  display: grid;
  grid-template-columns: repeat(3, minmax(0, 1fr));
  gap: 12px;
}

.platform-list div {
  min-height: 172px;
  display: grid;
  align-content: space-between;
  gap: 18px;
  padding: 22px;
  border: 1px solid rgba(17, 158, 146, 0.2);
  border-radius: var(--radius);
  background: rgba(251, 252, 248, 0.72);
}

.platform-list strong {
  font-size: 1.08rem;
}

.platform-list span {
  color: var(--muted);
  font-size: 0.96rem;
}

.pi-feature {
  display: grid;
  grid-template-columns: 280px minmax(0, 1fr);
  gap: 30px;
  align-items: start;
  padding: 24px;
  border: 1px solid var(--line);
  border-radius: var(--radius);
  background: var(--surface);
  box-shadow: 0 18px 48px rgba(34, 73, 65, 0.08);
}

.pi-feature img {
  width: 100%;
  aspect-ratio: 262 / 300;
  object-fit: cover;
  border-radius: var(--radius);
}

.pi-feature div {
  display: grid;
  gap: 14px;
}

.role-label {
  color: var(--accent-dark);
  font-weight: 800;
}

.text-link {
  color: var(--accent-dark);
  font-weight: 750;
  text-decoration: underline;
  text-decoration-thickness: 1px;
  text-underline-offset: 4px;
}

.people-grid {
  display: grid;
  grid-template-columns: repeat(4, minmax(0, 1fr));
  gap: 14px;
  margin-top: 18px;
}

.people-directory {
  display: grid;
  grid-template-columns: repeat(4, minmax(0, 1fr));
  gap: 16px;
  margin-top: 18px;
  align-items: start;
}

.person-card {
  min-width: 0;
  display: grid;
  gap: 10px;
  padding: 12px;
  border: 1px solid var(--line);
  border-radius: var(--radius);
  background: var(--surface);
}

.person-card img {
  width: 100%;
  aspect-ratio: 262 / 300;
  object-fit: cover;
  border-radius: calc(var(--radius) - 2px);
  background: var(--surface-strong);
}

.person-card h3 {
  font-size: 1rem;
}

.person-card p {
  font-size: 0.92rem;
}

.person-details {
  padding: 0;
  overflow: hidden;
}

.person-details summary {
  position: relative;
  display: grid;
  gap: 10px;
  min-height: 100%;
  padding: 12px 12px 44px;
  cursor: pointer;
  list-style: none;
}

.person-details summary::-webkit-details-marker {
  display: none;
}

.person-details summary::after {
  content: "+";
  position: absolute;
  right: 12px;
  bottom: 12px;
  width: 28px;
  height: 28px;
  display: grid;
  place-items: center;
  border: 1px solid rgba(17, 158, 146, 0.22);
  border-radius: 999px;
  color: var(--accent-dark);
  background: var(--accent-soft);
  font-weight: 800;
  line-height: 1;
}

.person-details[open] {
  box-shadow: 0 18px 48px rgba(34, 73, 65, 0.08);
}

.person-details[open] summary::after {
  content: "-";
}

.person-details-body {
  padding: 0 12px 16px;
}

.person-details-body p {
  padding-top: 14px;
  border-top: 1px solid var(--line);
  font-size: 0.94rem;
  line-height: 1.62;
}

.publication-section {
  padding: 100px 0;
  background: var(--surface);
  border-top: 1px solid var(--line);
  border-bottom: 1px solid var(--line);
}

.publication-page {
  padding-top: 84px;
  padding-bottom: 100px;
}

.publication-list {
  display: grid;
  gap: 0;
  border-top: 1px solid var(--line);
}

.publication-key {
  margin-bottom: 22px;
  color: var(--muted);
  font-size: 0.98rem;
}

.publication-row {
  display: grid;
  grid-template-columns: 180px minmax(0, 1fr);
  gap: 42px;
  padding: 28px 0;
  border-bottom: 1px solid var(--line);
}

.publication-meta {
  display: grid;
  align-content: start;
  gap: 8px;
}

.publication-meta span {
  color: var(--accent-dark);
  font-weight: 850;
}

.publication-meta small {
  color: var(--muted);
  font-size: 0.94rem;
}

.publication-body {
  display: grid;
  gap: 12px;
}

.publication-body h2 {
  max-width: 860px;
}

.publication-body a {
  text-decoration: none;
}

.publication-body a:hover,
.publication-body a:focus-visible {
  color: var(--accent-dark);
  text-decoration: underline;
  text-decoration-thickness: 1px;
  text-underline-offset: 4px;
}

.publication-note {
  margin-top: 28px;
  color: var(--muted);
}

.author-mark {
  color: var(--accent-dark);
  font-weight: 850;
  text-decoration: underline;
  text-decoration-thickness: 1px;
  text-underline-offset: 3px;
}

.publication-track {
  display: grid;
  grid-auto-flow: column;
  grid-auto-columns: minmax(320px, 420px);
  gap: 14px;
  overflow-x: auto;
  padding-bottom: 12px;
  scroll-snap-type: x mandatory;
}

.publication-card {
  min-height: 360px;
  display: grid;
  align-content: start;
  gap: 14px;
  padding: 24px;
  border: 1px solid var(--line);
  border-radius: var(--radius);
  background: var(--paper);
  scroll-snap-align: start;
}

.publication-card span {
  color: var(--accent-dark);
  font-size: 0.9rem;
  font-weight: 800;
}

.publication-card p {
  font-size: 0.96rem;
}

.contact-card {
  display: grid;
  grid-template-columns: minmax(0, 0.9fr) minmax(340px, 1fr);
  gap: 34px;
  padding: 36px;
  border-radius: var(--radius);
  background:
    linear-gradient(135deg, rgba(223, 243, 237, 0.94), rgba(251, 252, 248, 0.98)),
    var(--surface);
  border: 1px solid rgba(17, 158, 146, 0.22);
}

.contact-details {
  display: grid;
  gap: 14px;
}

.contact-card address {
  grid-column: 1 / -1;
  margin: 0;
  padding-top: 24px;
  border-top: 1px solid rgba(17, 158, 146, 0.22);
  font-style: normal;
}

.site-footer {
  padding: 34px 0;
}

.footer-grid {
  display: flex;
  justify-content: space-between;
  gap: 24px;
  color: var(--muted);
}

.footer-grid strong {
  color: var(--ink);
}

.reveal {
  opacity: 0;
  transform: translateY(18px);
  transition: opacity 520ms ease, transform 520ms ease;
}

.reveal.is-visible {
  opacity: 1;
  transform: translateY(0);
}

@media (max-width: 1080px) {
  .hero-grid {
    grid-template-columns: 1fr;
    gap: 26px;
  }

  .hero-brief {
    max-width: 720px;
  }

  .hero h1 {
    max-width: 760px;
    font-size: 3.7rem;
  }

  .intro-grid,
  .platform-grid,
  .research-grid {
    grid-template-columns: 1fr;
  }

  .research-row {
    grid-template-columns: 320px minmax(0, 1fr);
    gap: 22px;
  }

  .research-card-large {
    grid-row: auto;
  }

  .platform-list,
  .people-grid {
    grid-template-columns: repeat(2, minmax(0, 1fr));
  }

  .people-directory {
    grid-template-columns: repeat(3, minmax(0, 1fr));
  }
}

@media (max-width: 920px) {
  .people-directory {
    grid-template-columns: repeat(2, minmax(0, 1fr));
  }
}

@media (max-width: 760px) {
  .section-shell {
    width: min(100% - 28px, 1180px);
  }

  .site-header {
    grid-template-columns: auto auto;
    padding: 0 14px;
  }

  .menu-button {
    display: inline-flex;
    align-items: center;
  }

  .site-nav {
    position: fixed;
    top: var(--header-height);
    left: 14px;
    right: 14px;
    display: none;
    grid-template-columns: 1fr;
    padding: 12px;
    border: 1px solid var(--line);
    border-radius: var(--radius);
    background: var(--surface);
    box-shadow: var(--shadow);
  }

  .site-nav.is-open {
    display: grid;
  }

  .site-nav a {
    justify-content: center;
  }

  .hero {
    min-height: auto;
    background:
      linear-gradient(180deg, rgba(247, 249, 243, 0.98) 0%, rgba(247, 249, 243, 0.86) 56%, rgba(247, 249, 243, 0.66) 100%),
      url("assets/images/hero-evo-devo-background.png") 62% center / cover no-repeat;
  }

  .hero-grid {
    padding-top: 56px;
    padding-bottom: 72px;
  }

  .hero h1 {
    max-width: 360px;
    font-size: 2.65rem;
    line-height: 1;
  }

  .hero-brief {
    padding: 20px;
    background: rgba(251, 252, 248, 0.5);
  }

  .hero-brief p,
  .section-heading p,
  .platform-grid p,
  .contact-card > div > p {
    font-size: 1rem;
  }

  .intro-grid,
  .research-section,
  .people-section,
  .contact-section,
  .platform-section,
  .publication-section,
  .publication-page {
    padding-top: 64px;
    padding-bottom: 64px;
  }

  .intro-grid,
  .platform-grid,
  .contact-card,
  .pi-feature,
  .research-row,
  .publication-row {
    gap: 24px;
  }

  .intro-lede,
  .section-heading h2,
  .platform-grid h2,
  .contact-card h2,
  .page-hero h1 {
    font-size: 2.25rem;
  }

  .platform-list,
  .people-grid,
  .people-directory,
  .pi-feature,
  .research-row,
  .publication-row,
  .contact-card {
    grid-template-columns: 1fr;
  }

  .page-hero {
    padding: 70px 0 58px;
  }

  .pi-feature {
    padding: 16px;
  }

  .pi-feature img {
    max-width: 320px;
  }

  .publication-track {
    grid-auto-columns: minmax(280px, 86vw);
  }

  .publication-row {
    padding: 24px 0;
  }

  .publication-meta {
    grid-template-columns: auto auto;
    justify-content: start;
    align-items: baseline;
    gap: 12px;
  }

  .footer-grid {
    display: grid;
  }
}

@media (prefers-reduced-motion: reduce) {
  html {
    scroll-behavior: auto;
  }

  *,
  *::before,
  *::after {
    transition-duration: 0.01ms !important;
    animation-duration: 0.01ms !important;
    animation-iteration-count: 1 !important;
  }

  .reveal {
    opacity: 1;
    transform: none;
  }
}